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      "infons": {
        "treatment_title": "Omanitherium dhofarense SEIFFERT 2012", 
        "zenodo-doi": "10.2478/if-2017-0017", 
        "treatment-bank-uri": "http://treatment.plazi.org/id/9C33879B726EFFDCFC16FE60FBB53F6B", 
        "article-title": "Large Mammals From The Rupelian Of Oman - Recent Finds", 
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          "text": "Omanitherium dhofarense SEIFFERT 2012", 
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          }, 
          "text": "D e s c r i p t i o n. The fossils attributed to Omanitherium comprise 21 specimens out of 89 catalogue entries in the 2017 field catalogue (Tab. 2, Appendix 2), indicating that among the large mammals, this genus is dominant.", 
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                "evidence_code": "ECO_0000366", 
                "provenance": "SIBILS", 
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              "text": "fossils", 
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                "preferred_term": "fossils", 
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              "text": "fossils", 
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              "text": "mammals", 
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              "infons": {
                "type": "agriculture", 
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                "preferred_term": "mammals", 
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            "sentence_length": 137, 
            "offset": 263
          }, 
          "text": "Many of the specimens are broken pieces of mandibles and maxillae, and limb bone fragments, which yield little morphological information.", 
          "annotations": [
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              "id": "7", 
              "infons": {
                "type": "mesh", 
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                "concept_id": "D005121", 
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              "text": "limb", 
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                  "offset": 71, 
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              "id": "8", 
              "infons": {
                "type": "mesh", 
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                "version": "v2025", 
                "concept_id": "D008437", 
                "preferred_term": "Maxilla", 
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              "text": "maxillae", 
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        {
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            "sentence_number": 4, 
            "sentence_length": 37, 
            "offset": 400
          }, 
          "text": "There are also many enamel fragments.", 
          "annotations": []
        }, 
        {
          "infons": {
            "field": "text", 
            "sentence_number": 5, 
            "sentence_length": 272, 
            "offset": 437
          }, 
          "text": "The following descriptions deal only with the more complete dental elements, comprising some rather complete lower incisors, and a reasonable sample of upper cheek teeth, which were poorly represented in previously available samples (Seiffert et al. 2012, Pickford 2015b).", 
          "annotations": [
            {
              "id": "9", 
              "infons": {
                "type": "mesh", 
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                "concept_id": "D002610", 
                "preferred_term": "Cheek", 
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                "evidence_code": "ECO_0000366", 
                "provenance": "SIBILS", 
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              "text": "cheek", 
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              "text": "incisors", 
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              "infons": {
                "type": "agriculture", 
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                "preferred_term": "teeth", 
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                "provenance": "SIBILS", 
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              "text": "teeth", 
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            "sentence_length": 70, 
            "offset": 709
          }, 
          "text": "ONHM TN 2017-19 is a damaged left i/1 with apical wear (Text-fig. 10).", 
          "annotations": []
        }, 
        {
          "infons": {
            "field": "text", 
            "sentence_number": 7, 
            "sentence_length": 51, 
            "offset": 779
          }, 
          "text": "The section of the tooth is a compressed rectangle.", 
          "annotations": [
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              "id": "12", 
              "infons": {
                "type": "plant", 
                "concept_source": "po", 
                "version": "04-18-2024", 
                "concept_id": "PO:0025515", 
                "preferred_term": "phyllome lamina tooth", 
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                "evidence_code": "ECO_0000366", 
                "provenance": "SIBILS", 
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                "attributes": null
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              "text": "tooth", 
              "locations": [
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            "sentence_length": 170, 
            "offset": 830
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          "text": "The labial surface is almost complete and shows a shallow groove near the apex, positioned about one third of the distance from the mesial edge towards the distal margin.", 
          "annotations": [
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          "text": "The labial surface is shallowly convex, almost flat; the mesial surface is at right angles to the labial one and measures about 8 mm from labial to lingual.", 
          "annotations": [
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          "text": "The lingual surface is damaged, but remnants of enamel are preserved along the distal half, which shows that the lingual surface was originally covered in enamel, which is appreciably thinner than that on the labial side.", 
          "annotations": [
            {
              "id": "16", 
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        {
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          "text": "The distal edge is damaged.", 
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        {
          "infons": {
            "field": "text", 
            "sentence_number": 12, 
            "sentence_length": 120, 
            "offset": 1404
          }, 
          "text": "The cervix is distinct labially, and rises towards the apex on the mesial side and then extends across the lingual side.", 
          "annotations": []
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        {
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            "sentence_number": 13, 
            "sentence_length": 88, 
            "offset": 1524
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          "text": "The apex is somewhat damaged, so the exact shape of the wear facet cannot be determined.", 
          "annotations": []
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        {
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            "sentence_number": 14, 
            "sentence_length": 107, 
            "offset": 1612
          }, 
          "text": "ONHM TN 2017-51 is a complete, lightly worn right i/1 crown with part of the root preserved (Text-fig. 11).", 
          "annotations": [
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              "id": "18", 
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                "type": "species", 
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                "version": "v.2025-02-11", 
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                "evidence_code": "ECO_0000366", 
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                "concept_id": "c_16172", 
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                "evidence_code": "ECO_0000366", 
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              "text": "crown", 
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            "offset": 1719
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          "text": "The tooth has a rectangular section, much broader mesiodistally than the labio-lingual diameter.", 
          "annotations": [
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          "text": "The labial surface has a subtle groove that extends from the apex towards the root, fading out as it goes.", 
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        }, 
        {
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            "offset": 1921
          }, 
          "text": "This groove is positioned about 1/3 of the distance between the mesial and distal margins of the crown.", 
          "annotations": [
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              "id": "27", 
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                "concept_source": "agrovoc", 
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              "text": "crown", 
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        }, 
        {
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          }, 
          "text": "The mesial side of the tooth is almost flat, with a complete cover of thin enamel.", 
          "annotations": [
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              "text": "tooth", 
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        {
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          "text": "The distal margin of the tooth is curved apically, but for most of its height it is parallel to the mesial margin.", 
          "annotations": [
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          "text": "The mesial side of the tooth is flat and at right angles to the labial side.", 
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          "text": "The wear facet is straight and at right angles to the long axis of the crown.", 
          "annotations": [
            {
              "id": "32", 
              "infons": {
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            "offset": 2373
          }, 
          "text": "ONHM TN 2017-43 is a left i/2 with a wear facet at its apex.", 
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        }, 
        {
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            "offset": 2433
          }, 
          "text": "The section is ovoid, such that the mesio-distal diameter is slightly greater than the labio-lingual diameter.", 
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            "offset": 2543
          }, 
          "text": "The labial surface is uniformly curved, but the lingual side is more angular.", 
          "annotations": [
            {
              "id": "36", 
              "infons": {
                "type": "conceptual_entity", 
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            "offset": 2620
          }, 
          "text": "The cervix is far from the apex on the labial side, but rises a long way towards the apex mesially, before descending rootwards again.", 
          "annotations": []
        }, 
        {
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          "text": "The wear facet is flat and at right angles to the long axis of the tooth.", 
          "annotations": [
            {
              "id": "37", 
              "infons": {
                "type": "plant", 
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            "sentence_length": 122, 
            "offset": 2827
          }, 
          "text": "ONHM TN 2017-50 is the distal lophid of an unworn lower molar, probably m/2 on the basis of its dimensions (Text-fig. 12).", 
          "annotations": [
            {
              "id": "38", 
              "infons": {
                "type": "mesh", 
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                "concept_id": "D008963", 
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            "offset": 2949
          }, 
          "text": "The crest of the lophid has a mamellated appearance, with no central sulcus.", 
          "annotations": [
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              "infons": {
                "type": "gene", 
                "concept_source": "uniprot_swissprot", 
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                "preferred_term": "Stannous Fluoride", 
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            "sentence_length": 76, 
            "offset": 3025
          }, 
          "text": "The precristids are welldeveloped, in particular the one on the buccal cusp.", 
          "annotations": [
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          }, 
          "text": "The distal cingulum is broken off, but the posterior accessory tubercle is preserved, and lies in line with the pre-cristid of the buccal cusp, as in other specimens of the species (Textfig. 13).", 
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          "text": "ONHM TN 2017-16 is an unworn left P3/ crown.", 
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              "infons": {
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            "offset": 3340
          }, 
          "text": "It has a prominent protocone, which is separated by a sulcus from the buccal cusp, which is comprised of closely fused paracone and metacone.", 
          "annotations": [
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          }, 
          "text": "The parastyle is bordered on either side by a cingulum, which is prominent near the parastyle, but fades out buccally and lingually.", 
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        {
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          "text": "The distal cingulum is prominent in the centre-line of the tooth, but diminishes buccally and lingually.", 
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          "text": "The pre-protocrista reaches mesiobuccally towards the pre-paracrista, thereby forming a lophlike wall with a shallow notch between the two cristae.", 
          "annotations": [
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              }, 
              "text": "forming", 
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        }, 
        {
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            "offset": 3864
          }, 
          "text": "ONHM TN 2017-17 is an unworn right P4/ found close to the left P3/.", 
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        }, 
        {
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            "offset": 3931
          }, 
          "text": "It is slightly larger than the P3/, the loph is better formed and the central sulcus is broader than that in the P3/.", 
          "annotations": [
            {
              "id": "54", 
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          "text": "The protocone is large, and the paracone and metacone are separated at their apices, but solidly fused at their bases.", 
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        }, 
        {
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            "offset": 4166
          }, 
          "text": "The crests of the tooth are comparable to those in the P3/, but show stronger beading along their edges.", 
          "annotations": [
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              "infons": {
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            "offset": 4270
          }, 
          "text": "The same can be said about the beading of the cingula.", 
          "annotations": [
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              "text": "cingula", 
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          "text": "Catalogue n\u00b0 Tooth Mesio-distal length Bucco-lingual breadth Crown height ONHM TN 2017-51 Right i/1 31.0 22.7 49+ ONHM TN 2017-19 Left i/1 29.4 \u2013 45+ ONHM TN 2017-43 Left i/2 27.6 20.7 50+ ONHM TN 2017-85 Right i/2 32.0 23.0 56.0 ONHM TN 2017-50 Left m/2 \u2013 37.3 \u2013 ONHM TN 2017-16 Left P3/ 22.5 24.0 \u2013 ONHM TN 2017-17 Right P4/ 22.3 29.0 \u2013 ONHM TN 2017-44 Right P4/ 24.4 30.7 \u2013 ONHM TN 2017-18 Right M1/ 31.2 \u2013 \u2013 ONHM TN 2017-79 Left M3/ 43.7 38.3 \u2013\nONHM TN 2017-44 is a lightly worn right P4/, similar in construction to ONHM TN 2017-17, but slightly larger (Text-fig. 14).", 
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          "text": "The apices of the loph and the cingula are lightly worn, to the stage where the beading has been removed.", 
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          "text": "ONHM TN 2017-18 is a lightly worn right M1/, lacking the buccal part of the crown (Text-fig. 14).", 
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          "text": "It was found close to the left P3/.", 
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          }, 
          "text": "The protocone and paracone form a clear loph, as do the hypocone and metacone behind, separated from each other by a broad central valley.", 
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          "text": "There is a narrow mesial cingulum, and a broader distal one.", 
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          "text": "The mesial edge of each loph is straight, but the distal side is concave, due to the way that the post-cristae are arranged.", 
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          "text": "The mesial cingulum is broken off, but the distal one is preserved, and shows a beaded apex.", 
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          "text": "The distal cingulum blends into the post-metacrista.", 
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          "text": "This is due to the fact that not only is the hypocone more distally positioned relative to the metacone than it is on the anterior molars, but in addition, the post-hypocrista is more voluminous than the post-metacrista.", 
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          "text": "As a result of this arrangement of the main cusps, the distal cingulum is oblique to the long axis of the tooth.", 
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          }, 
          "text": "In other respects, the M3/ is similar to the other molars.", 
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            "sentence_length": 148, 
            "offset": 6320
          }, 
          "text": "D i s c u s s i o n. The first mentions of the presence of a lophodont proboscidean at Thaytiniti were by Thomas et al. (1989, 1999), summarised as?", 
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        }, 
        {
          "infons": {
            "field": "text", 
            "sentence_number": 58, 
            "sentence_length": 22, 
            "offset": 6468
          }, 
          "text": "Barytherium sp. indet.", 
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              "text": "Barytherium", 
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            "sentence_length": 5, 
            "offset": 6490
          }, 
          "text": "(\u201ccf.", 
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        }, 
        {
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            "field": "text", 
            "sentence_number": 60, 
            "sentence_length": 108, 
            "offset": 6495
          }, 
          "text": "Barytherioidea\u201d) by Sanders et al. (2010b), and now known as Omanitherium dhofarense SEIFFERT et al., 2012 .", 
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            "sentence_length": 167, 
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          }, 
          "text": "When Omanitherium was described for the first time, Seiffert et al. (2012) reconstructed the mandible with only two incisors, separated from each other by a broad gap.", 
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          "text": "In contrast, Pickford (2015b) thought that there would have been a pair of central incisors infilling the gap.", 
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          "text": "The new fossils from the type locality and nearby sites indicate that Omanitherium did indeed possess four lower incisors, but instead of the central lower incisors being relatively low crowned as originally thought, they are taller.", 
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